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Unlikely to have "hundreds of volts"

Although neither of us can be sure, it is probable that the DC voltage on the line between the output of the DAC and the input of whatever is less than 50V. Because the output stage is likely to be a cathode follower, which means that the output is taken from the cathode of the last tube. The VDC between the cathode and ground is typically way less than the plate voltage. However, I share your concern in a general sense.

One other thing that I wonder about: The OP says that the input capacitor is 10uF value. I wonder why the value would be so large. This suggests that the input impedance, largely determined by the resistance between the input jack and ground, is rather low. If one knew more about THAT circuit, one might justify removing the input capacitor, rather than the output capacitor from the DAC, especially if the input capacitor is an electrolytic. But don't, without knowing what's up with the input stage of the driven component. It's possible that the input capacitor is blocking grid bias voltage from the upstream component, in which case it's not safe to remove it (unless and only possibly if you re-install the output cap).

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